Stromules are thin, tubular extensions that form on the surface of plastids inside plant cells. These structures contain plastid stroma and can change shape and length quickly, projecting away from organelles such as chloroplasts.

Historical observations

Descriptions of plastid protrusions date back to the late 19th and early 20th centuries, with reports from 1888 and 1908 noting outgrowths and connections between plastids. Interest in these features increased after they were brought back to attention in 1997, when modern imaging made their presence and dynamics clearer.

Where they occur

Stromules extend from many types of plastids and have been recorded across a variety of flowering plants. Examples include Arabidopsis thaliana, wheat, rice and tomato.

Function and current research

Researchers continue to investigate stromules. Their transient, often branching form suggests roles in increasing plastid surface area, facilitating molecular exchange between plastids and other cell compartments, or in signaling, but none of these functions has been definitively established.